germ, may, by acting on these, superinduce further complications; upon these still higher and more numerous ones; and so on continually: each organ as it is developed serving, by its actions and reactions on the rest, to initiate new complexities. The first pulsations of the fœtal heart must simultaneously aid the unfolding of every part. The growth of each tissue, by taking from the blood special proportions of elements, must modify the constitution of the blood; and so must modify the nutrition of all the other tissues. The heart's action, implying as it does a certain waste, necessitates an addition to the blood of effete matters, which must influence the rest of the system, and perhaps, as some think, cause the formation of excretory organs. The nervous connexions established among the viscera must further multiply their mutual influences; and so continually. Still stronger becomes the probability of this view when we call to mind the fact, that the same germ may be evolved into different forms according to circumstances. Thus, during its earlier stages, every embryo is sexless – becomes either male or female as the balance of forces acting on it determines. Again, it is a well-established fact that the larva of a working-bee will develop into a queen-bee, if before it is too late, its food be changed to that on which the larvæ of queen-bees are fed. All which instances suggest that the proximate cause of each advance in embryonic complication is the action of incident forces upon the complication previously existing. Indeed, we may find a priori reason to think that the evolution proceeds after this manner. For since no germ, animal or vegetal, contains the slightest rudiment or indication of the future organism – since the microscope has shown us that the first process set up in every fertilized germ, is a process of repeated spontaneous fissions ending in the production of a mass of cells, not one of which exhibits any special character; there seems no alternative but to suppose that the partial organization at any moment existing in a growing embryo, is transformed by the agencies acting upon it into the succeeding phase of organization, and this into the next, until, through ever-increasing complexities, the ultimate form is reached. Not indeed that we can thus really explain the production of any plant or animal. We are still in the dark respecting those mysterious properties in virtue of which the germ, when subject to fit influences, undergoes the special changes that begin the series of transformations. All we aim to show, is, that given a germ possessing those particular proclivities distinguishing the species to which it belongs, and the evolution of an organism from it, probably depends on that multiplication of effects which we have seen to be the cause of progress in general, so far as we have yet traced it.
When, leaving the development of single plants and animals, we pass to that of the Earth's flora and fauna, the course of our argument again becomes clear and simple. Though, as was admitted in the first part of this article, the fragmentary facts Paleontology has accumulated, do not clearly warrant us in saying that, in the lapse of geologic time, there have been evolved more heterogeneous organisms, and more heterogeneous assemblages of organisms, yet we shall now see that there must ever have been a tendency towards these results. We shall find that the production of many effects by one cause, which as already shown, has been all along increasing the physical heterogeneity of the Earth, has further involved an increasing heterogeneity in its flora and fauna, individually and collectively. An illustration will make this clear. Suppose that by a series of upheavals, occurring, as they are now known to do, at long intervals, the East Indian Archipelago were to be, step by step, raised into a continent, and a chain of mountains formed along the axis of elevation. By the first of these upheavals, the plants and animals inhabiting Borneo, Sumatra, New Guinea, and the rest, would be subjected to slightly modified sets of conditions. The climate in general would be altered in temperature, in humidity, and in its periodical variations; while the local differences would be multiplied. These modifications would affect, perhaps inappreciably, the entire flora and fauna of the region. The change of level would produce additional modifications: varying in different species, and also in different members of the same species, according to their distance from the axis of elevation. Plants, growing only on the sea-shore in special localities, might become extinct. Others, living only in swamps of a certain humidity, would, if they survived at all, probably undergo visible changes of appearance. While still greater alterations would occur in the plants gradually spreading over the lands newly raised above the sea. The animals and insects living on these modified plants, would themselves be in some degree modified by change of food, as well as by change of climate; and the modification would be more marked where, from the dwindling or disappearance of one kind of plant, an allied kind was eaten. In the lapse of the many generations arising before the next upheaval, the sensible or insensible alterations thus produced in each species would become organized – there would be a more or less complete adaptation to the new conditions. The next upheaval would superinduce further organic changes, implying wider divergences from the primary forms; and so repeatedly. But now let it be observed that the revolution thus resulting would not be a substitution of a thousand more or less modified species for the thousand original species; but in place of the thousand original species there would arise several thousand species, or varieties, or changed forms. Each species being distributed over an area of some extent, and tending continually to colonize the new area exposed, its different members would be subject to different sets of changes. Plants and animals spreading towards the equator would not be affected in the same way as others spreading from it. Those spreading towards the new shores would undergo changes unlike the changes undergone by those spreading into the mountains. Thus, each original race of organisms, would become the root from which diverged several races differing more or less from it and from each other; and while some of these might subsequently disappear, probably more than one would survive in the next geologic period: the very dispersion itself increasing the chances of survival. Not only would there be certain modifications thus caused by change of physical conditions and food, but also in some cases other modifications caused by change of habit. The fauna of each island, peopling, step by step, the newly-raised tracts, would eventually come in contact with the faunas of other islands; and some members of these other faunas would be unlike any creatures before seen. Herbivores meeting with new beasts of prey, would, in some cases, be led into modes of defence or escape differing from those previously used; and simultaneously the beasts of prey would modify their modes of pursuit and attack. We know that when circumstances demand it, such changes of habit do take place in animals; and we know that if the new habits become the dominant ones, they must eventually in some degree alter the organization. Observe now, however, a further consequence. There must arise not simply a tendency towards the differentiation of each race of organisms into several races; but also a tendency to the occasional production of a somewhat higher organism. Taken in the mass these divergent varieties which have been caused by fresh physical conditions and habits of life, will exhibit changes quite indefinite in kind and degree; and changes that do not necessarily constitute an advance. Probably in most cases the modified type will be neither more nor less heterogeneous than the original one. In some cases the habits of life adopted being simpler than before, a less heterogeneous structure will result: there will be a retrogradation. But it must now and then occur, that some division of a species, falling into circumstances which give it rather more complex experiences, and demand actions somewhat more involved, will have certain of its organs further differentiated in proportionately small degrees, – will become slightly more heterogeneous. Thus, in the natural course of things, there will from time to time arise an increased heterogeneity both of the Earth's flora and fauna, and of individual races included in them. Omitting detailed explanations, and allowing for the qualifications which cannot here be specified, we think it is clear that geological mutations have all along tended to complicate the forms of life, whether regarded separately or collectively. The same causes which have led to the evolution of the Earth's crust from the simple into the complex, have simultaneously led to a parallel evolution of the Life upon its surface. In this case, as in previous ones, we see that the transformation of the homogeneous into the heterogeneous is consequent upon the universal principle, that every active force produces more than one change.
The deduction here drawn from the established truths of geology and the general laws of life, gains immensely in weight on finding it to be in harmony with an induction drawn from direct experience. Just that divergence of many races from one race, which we inferred must have been continually occurring during geologic time, we know to have occurred during the pre-historic and historic periods, in man and domestic animals. And just that multiplication of effects which we concluded must have produced the first, we see has produced the last. Single causes,