Spencer Herbert

The Principles of Biology, Volume 1 (of 2)

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to the character and derivation of the cells (muscle-corpuscles) the musculature is divided into epithelial (epithelial muscle-cells, primary bundles) and connective-tissue muscle cells (contractile fibre-cells).

      "The physiological character of nervous tissue rests upon the transmission of sensory stimuli and voluntary impulses, and upon the co-ordination of these into unified psychic activity.

      "The conduction takes place by means of nerve-fibres (non-medullated and medullated fibrils and bundles of fibrils); the co-ordination of stimuli by means of ganglion-cells (bipolar, multipolar ganglion-cells)." (General Principles of Zoology, pp. 117-8.)

      But now concerning cells out of which, variously modified, obscured, and sometimes obliterated, tissues are formed, we have to note a fact of much significance. Along with the cell-doctrine as at first held, when attention was given to the cell itself rather than to its contents, there went the belief that each of these morphological units is structurally separate from its neighbours. But since establishment of the modern view that the essential element is the contained protoplasm, histologists have discovered that there are protoplasmic connexions between the contents of adjacent cells. Though cursorily observed at earlier dates, it was not until some twenty years ago that in plant-tissues these were clearly shown to pass through openings in the cell-walls. It is said that in some cases the openings are made, and the junctions established, by a secondary process; but the implication is that usually these living links are left between multiplying protoplasts; so that from the outset the protoplasm pervading the whole plant maintains its continuity. More recently sundry zoologists have alleged that a like continuity exists in animals. Especially has this been maintained by Mr. Adam Sedgwick. Numerous observations made on developing ova of fishes have led him to assert that in no case do the multiplying cells so-called – blastomeres and their progeny – become entirely separate. Their fission is in all cases incomplete. A like continuity has been found in the embryos of many Arthropods, and more recently in the segmenting eggs and blastulæ of Echinoderms. The syncytium thus formed is held by Mr. Sedgwick to be maintained in adult life, and in this belief he is in agreement with sundry others. Bridges of protoplasm have been seen between epithelium-cells, and it is maintained that cartilage-cells, connective tissue cells, the cells forming muscle-fibres, as well as nerve-cells, have protoplasmic unions. Nay, some even assert that an ovum preserves a protoplasmic connexion with the matrix in which it develops.

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      1

      Gross misrepresentations of this statement, which have been from time to time made, oblige me, much against my will, to add here an explanation of it. The last of these perversions, uttered in a lecture delivered at Belfast by the Rev. Professor Watts, D.D., is reported in the Belfast Witness of December 18, 1874; just while a third impression of this work is being printed from the plates. The report commences as follows: – "Dr. Watts, after showing that on his own confession Spencer was indebted for his facts to Huxley and Hooker, who," &c., &c.

      Wishing in this, as in other cases, to acknowledge indebtedness when conscious of it, I introduced the words referred to, in recognition of the fact that I had repeatedly questioned the distinguished specialists named, on matters beyond my knowledge, which were not dealt with in the books at my command. Forgetting the habits of antagonists, and e

1

Gross misrepresentations of this statement, which have been from time to time made, oblige me, much against my will, to add here an explanation of it. The last of these perversions, uttered in a lecture delivered at Belfast by the Rev. Professor Watts, D.D., is reported in the Belfast Witness of December 18, 1874; just while a third impression of this work is being printed from the plates. The report commences as follows: – "Dr. Watts, after showing that on his own confession Spencer was indebted for his facts to Huxley and Hooker, who," &c., &c.

Wishing in this, as in other cases, to acknowledge indebtedness when conscious of it, I introduced the words referred to, in recognition of the fact that I had repeatedly questioned the distinguished specialists named, on matters beyond my knowledge, which were not dealt with in the books at my command. Forgetting the habits of antagonists, and especially theological antagonists, it never occurred to me that my expression of thanks to my friends for "information where my own was deficient," would be turned into the sweeping statement that I was indebted to them for my facts.

Had Professor Watts looked at the preface to the second volume (the two having been published separately, as the prefaces imply), he would have seen a second expression of my indebtedness "for their valuable criticisms, and for the trouble they have taken in checking the numerous statements of fact on which the arguments proceed" – no further indebtedness being named. A moment's comparison of the two volumes in respect of their accumulations of facts, would have shown him what kind of warrant there was for his interpretation.

Doubtless the Rev. Professor was prompted to make this assertion by the desire to discredit the work he was attacking; and having so good an end in view, thought it needless to be particular about the means. In the art of dealing with the language of opponents, Dr. Watts might give lessons to Monsignor Capel and Archbishop Manning.

December 28th, 1874.

2

In this passage as originally written (in 1862) they were described as incondensible; since, though reduced to the density of liquids, they had not been liquefied.

3

Here and hereafter the word "atom" signifies a unit of something classed as an element, because thus far undecomposed by us. The word must not be supposed to mean that which its derivation implies. In all probability it is not a simple unit but a compound one.

4

The name hydro-carbons was here used when these pages were written, thirty-four years ago. It was the name then current. In this case, as in multitudinous other cases, the substitution of newer words and phrases for older ones, is somewhat misleading. Putting the thoughts of 1862 in the language of 1897 gives an illusive impression of recency.

5

It will perhaps seem strange to class oxygen as a crystalloid. But inasmuch as the crystalloids are distinguished from the colloids by their atomic simplicity, and inasmuch as sundry gases are reducible to a crystalline state, we are justified in so classing it.

6

The remark made by a critic to the effect that in a mammal higher temperature diminishes the rate of molecular change in the tissues, leads me to add that the exhalation I have alleged is prevented if the heat rises above the range of variation normal to the organism; since, then, unusually rapid pulsations with consequent inefficient propulsion of the blood, cause a diminished rate of circulation. To produce the effect referred to in the text, heat must be associated with dryness; for otherwise evaporation is not aided. General evidence supporting the statement I have made is furnished by the fact that the hot and dry air of the eastern deserts is extremely invigorating; by the fact that all the energetic and conquering races of men have come from the hot and dry regions marked on the maps as rainless; and by the fact that travellers in Africa comment on the contrast between the inhabitants of the hot and dry regions (relatively elevated) and those of the hot and moist regions: active and inert respectively.

7

The

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