before the Glacial period had begun to whiten the distant mountains of central and northern Europe, fell exhausted at last upon the shore of Terceira. There were no insects then for the poor bird to feed upon, so it died of starvation and weariness before the day was out; but a little earth that clung in a pellet to one of its feet contained the egg of a land-shell, while the prickly seed of a common Spanish plant was entangled among the winged feathers by its hooked awns. The egg hatched out, and became the parent of a large brood of minute snails, which, outliving the cold spell of the Ice Age, had developed into a very distinct type in the long period that intervened before the advent of man in the islands; while the seed sprang up on the natural manure heap afforded by the swallow's decaying body, and clinging to the valleys during the Glacial Age on the hill-tops, gave birth in due season to one of the most markedly indigenous of our Terceira plants.
Occasionally, too, very minute land-snails would arrive alive on the island after their long sea-voyage on bits of broken forest-trees—a circumstance which I would perhaps hesitate to mention in mere human society were it not that I have been credibly informed your own great naturalist, Darwin, tried the experiment himself with one of the biggest European land-molluscs, the great edible Roman snail, and found that it still lived on in vigorous style after immersion in sea-water for twenty days. Now, I myself observed that several of these bits of broken trees, torn down by floods in heavy storm time from the banks of Spanish or Portuguese rivers, reached my island in eight or ten days after leaving the mainland, and sometimes contained eggs of small land-snails. But as very long periods often passed without a single new species being introduced into the group, any kind that once managed to establish itself on any of the islands usually remained for ages undisturbed by new arrivals, and so had plenty of opportunity to adapt itself perfectly by natural selection to the new conditions. The consequence was, that out of some seventy land-snails now known in the islands, thirty-two had assumed distinct specific features before the advent of man, while thirty-seven (many of which, I think, I never noticed till the introduction of cultivated plants) are common to my group with Europe or with the other Atlantic islands. Most of these, I believe, came in with man and his disconcerting agriculture.
As to the pond and river snails, so far as I could observe, they mostly reached us later, being conveyed in the egg on the feet of stray waders or water-birds, which gradually peopled the island after the Glacial epoch.
Birds and all other flying creatures are now very abundant in all the islands; but I could tell you some curious and interesting facts, too, as to the mode of their arrival and the vicissitudes of their settlement. For example, during the age of the Forest Beds in Europe, a stray bullfinch was driven out to sea by a violent storm, and perched at last on a bush at Fayal. I wondered at first whether he would effect a settlement. But at that time no seeds or fruits fit for bullfinches to eat existed on the islands. Still, as it turned out, this particular bullfinch happened to have in his crop several undigested seeds of European plants exactly suited to the bullfinch taste; so when he died on the spot, these seeds, germinating abundantly, gave rise to a whole valleyful of appropriate plants for bullfinches to feed upon. Now, however, there was no bullfinch to eat them. For a long time, indeed, no other bullfinches arrived at my archipelago. Once, to be sure, a few hundred years later, a single cock bird did reach the island alone, much exhausted with his journey, and managed to pick up a living for himself off the seeds introduced by his unhappy predecessor. But as he had no mate, he died at last, as your lawyers would say, without issue.
It was a couple of hundred years or so more before I saw a third bullfinch—which didn't surprise me, for bullfinches are very woodland birds, and non-migratory into the bargain—so that they didn't often get blown seaward over the broad Atlantic. At the end of that time, however, I observed one morning a pair of finches, after a heavy storm, drying their poor battered wings upon a shrub in one of the islands. From this solitary pair a new race sprang up, which developed after a time, as I imagined they must, into a distinct species. These local bullfinches now form the only birds peculiar to the islands; and the reason is one well divined by one of your own great naturalists (to whom I mean before I end to make the amende honorable). In almost all other cases the birds kept getting reinforced from time to time by others of their kind blown out to sea accidentally—for only such species were likely to arrive there—and this kept up the purity of the original race, by ensuring a cross every now and again with the European community. But the bullfinches, being the merest casuals, never again to my knowledge were reinforced from the mainland, and so they have produced at last a special island type, exactly adapted to the peculiarities of their new habitat.
You see, there was hardly ever a big storm on land that didn't bring at least one or two new birds of some sort or other to the islands. Naturally, too, the newcomers landed always on the first shore they could sight; and so at the present day the greatest number of species is found on the two easternmost islands nearest the mainland, which have forty kinds of land-birds, while the central islands have but thirty-six, and the western only twenty-nine. It would have been quite different, of course, if the birds came mainly from America with the trade winds and the Gulf Stream, as I at first anticipated. In that case, there would have been most kinds in the westernmost islands, and fewest stragglers in the far eastern. But your own naturalists have rightly seen that the existing distribution necessarily implies the opposite explanation.
Birds, I early noticed, are always great carriers of fruit-seeds, because they eat the berries, but don't digest the hard little stones within. It was in that way, I fancy, that the Portugal laurel first came to my islands, because it has an edible fruit with a very hard seed; and the same reason must account for the presence of the myrtle, with its small blue berry; the laurustinus with its currant-like fruit; the elder-tree, the canary laurel, the local sweet-gale, and the peculiar juniper. Before these shrubs were introduced thus unconsciously by our feathered guests, there were no fruits on which berry-eating birds could live; but now they are the only native trees or large bushes on the islands—I mean the only ones not directly planted by you mischief-making men, who have entirely spoilt my nice little experiment.
It was much the same with the history of some among the birds themselves. Not a few birds of prey, for example, were driven to my little archipelago by stress of weather in its very early days; but they all perished for want of sufficient small quarry to make a living out of. As soon, however, as the islands had got well stocked with robins, black-caps, wrens, and wagtails, of European types—as soon as the chaffinches had established themselves on the seaward plains, and the canary had learnt to nest without fear among the Portugal laurels—then buzzards, long-eared owls, and common barn-owls, driven westward by tempests, began to pick up a decent living on all the islands, and have ever since been permanent residents, to the immense terror and discomfort of our smaller song-birds. Thus the older the archipelago got the less chance was there of local variation taking place to any large degree, because the balance of life each day grew more closely to resemble that which each species had left behind it in its native European or African mainland.
I said a little while ago we had no mammal in the islands. In that I was not quite strictly correct. I ought to have said, no terrestrial mammal. A little Spanish bat got blown to us once by a rough nor'easter, and took up its abode at once among the caves of our archipelago, where it hawks to this day after our flies and beetles. This seemed to me to show very conspicuously the advantage which winged animals have in the matter of cosmopolitan dispersion; for while it was quite impossible for rats, mice, or squirrels to cross the intervening belt of three hundred leagues of sea, their little winged relation, the flitter-mouse, made the journey across quite safely on his own leathery vans, and with no greater difficulty than a swallow or a wood-pigeon.
The insects of my archipelago tell very much the same story as the birds and the plants. Here, too, winged species have stood at a great advantage. To be sure, the earliest butterflies and bees that arrived in the fern-clad period were starved for want of honey; but as soon as the valleys began to be thickly tangled with composites, harebells, and sweet-scented myrtle bushes, these nectar-eating insects established themselves successfully, and kept their breed true by occasional crosses with fresh arrivals blown to sea afterwards. The development of the beetles I watched with far greater interest, as they assumed fresh forms much more rapidly under their new conditions of restricted food and limited enemies. Many kinds I observed which came originally from Europe, sometimes in the larval state, sometimes in the egg, and sometimes