cause of such change would be found in the diminution of the area covered by the forests.
The industry of man is said to have been so successful in the local extirpation of noxious or useless vegetables in China, that, with the exception of a few water plants in the rice grounds, it is sometimes impossible to find a single weed in an extensive district; and the late eminent agriculturist, Mr. Coke, is reported to have offered in vain a considerable reward for the detection of a weed in a large wheatfield on his estate in England. In these cases, however, there is no reason to suppose that diligent husbandry has done more than to eradicate the pests of agriculture within a comparatively limited area, and the cockle and the darnel will probably remain to plague the slovenly cultivator as long as the cereal grains continue to bless him.[56]
Origin of Domestic Plants.
One of the most important, and, at the same time, most difficult questions connected with our subject is: how far we are to regard our cereal grains, our esculent bulbs and roots, and the multiplied tree fruits of our gardens, as artificially modified and improved forms of wild, self-propagating vegetation. The narratives of botanical travellers have often announced the discovery of the original form and habitat of domesticated plants, and scientific journals have described the experiments by which the identity of particular wild and cultivated vegetables has been thought to be established. It is confidently affirmed that maize and the potato—which we must suppose to have been first cultivated at a much later period than the breadstuffs and most other esculent vegetables of Europe and the East—are found wild and self-propagating in Spanish America, though in forms not recognizable by the common observer as identical with the familiar corn and tuber of modern agriculture. It was lately asserted, upon what seemed very strong evidence, that the Ægilops ovata, a plant growing wild in Southern France, had been actually converted into common wheat; but, upon a repetition of the experiments, later observers have declared that the apparent change was only a case of temporary hybridation or fecundation by the pollen of true wheat, and that the grass alleged to be transformed into wheat could not be perpetuated as such from its own seed.
The very great modifications which cultivated plants are constantly undergoing under our eyes, and the numerous varieties and races which spring up among them, certainly countenance the doctrine, that every domesticated vegetable, however dependent upon human care for growth and propagation in its present form, may have been really derived, by a long succession of changes, from some wild plant not now much resembling it. But it is, in every case, a question of evidence. The only satisfactory proof that a given wild plant is identical with a given garden or field vegetable, is the test of experiment, the actual growing of the one from the seed of the other, or the conversion of the one into the other by transplantation and change of conditions. It is hardly contended that any of the cereals or other plants important as human aliment, or as objects of agricultural industry, exist and propagate themselves uncultivated in the same form and with the same properties as when sown and reared by human art.[57] In fact, the cases are rare where the identity of a wild with a domesticated plant is considered by the best authorities as conclusively established, and we are warranted in affirming of but few of the latter, as a historically known or experimentally proved fact, that they ever did exist, or could exist, independently of man.[58]
Organic Life as a Geological and Geographical Agency.
The quantitative value of organic life, as a geological agency, seems to be inversely as the volume of the individual organism; for nature supplies by numbers what is wanting in the bulk of the plant or animal out of whose remains or structures she forms strata covering whole provinces, and builds up from the depths of the sea large islands, if not continents. There are, it is true, near the mouths of the great Siberian rivers which empty themselves into the Polar Sea, drift islands composed, in an incredibly large proportion, of the bones and tusks of elephants, mastodons, and other huge pachyderms, and many extensive caves in various parts of the world are half filled with the skeletons of quadrupeds, sometimes lying loose in the earth, sometimes cemented together into an osseous breccia by a calcareous deposit or other binding material. These remains of large animals, though found in comparatively late formations, generally belong to extinct species, and their modern congeners or representatives do not exist in sufficient numbers to be of sensible importance in geology or in geography by the mere mass of their skeletons.[59] But the vegetable products found with them, and, in rare cases, in the stomachs of some of them, are those of yet extant plants; and besides this evidence, the recent discovery of works of human art, deposited in juxtaposition with fossil bones, and evidently at the same time and by the same agency which buried these latter—not to speak of alleged human bones found in the same strata—proves that the animals whose former existence they testify were contemporaneous with man, and possibly even extirpated by him.[60] I do not propose to enter upon the thorny question, whether the existing races of man are genealogically connected with these ancient types of humanity, and I advert to these facts only for the sake of the suggestion that man, in his earliest known stages of existence, was probably a destructive power upon the earth, though perhaps not so emphatically as his present representatives.
The larger wild animals are not now numerous enough in any one region to form extensive deposits by their remains; but they have, nevertheless, a certain geographical importance. If the myriads of large browsing and grazing quadrupeds which wander over the plains of Southern Africa—and the slaughter of which by thousands is the source of a ferocious pleasure and a brutal triumph to professedly civilized hunters—if the herds of the American bison, which are numbered by hundreds of thousands, do not produce visible changes in the forms of terrestrial surface, they have at least an immense influence on the growth and distribution of vegetable life, and, of course, indirectly upon all the physical conditions of soil and climate between which and vegetation a mutual interdependence exists.
The influence of wild quadrupeds upon vegetable life has been little studied, and not many facts bearing upon it have been recorded, but, so far as it is known, it appears to be conservative rather than pernicious.[61] Few if any of them depend for their subsistence on vegetable products obtainable only by the destruction of the plant, and they seem to confine their consumption almost exclusively to the annual harvest of leaf or twig, or at least of parts of the vegetable easily reproduced. If there are exceptions to this rule, they are in cases where the numbers of the animal are so proportioned to the abundance of the vegetable, that there is no danger of the extermination of the plant from the voracity of the quadruped, or of the extinction of the quadruped from the scarcity of the plant. In diet and natural wants the bison resembles the ox, the ibex and the chamois assimilate themselves to the goat and the sheep; but while the wild animal does not appear to be a destructive agency in the garden of nature, his domestic congeners are eminently so. This is partly from the change of habits resulting from domestication and association with man, partly from the fact that the number of reclaimed animals is not determined by the natural relation of demand and spontaneous supply which regulates the multiplication of wild creatures, but by the convenience of man, who is, in comparatively few things, amenable to the control of the merely physical arrangements of nature. When the domesticated animal escapes from human jurisdiction, as in the case of the ox, the horse, the goat, and perhaps the ass—which, so far as I know, are the only well-authenticated instances of the complete emancipation of household quadrupeds—he becomes again an unresisting subject of nature, and all his economy is governed by the same laws as that of his fellows which have never been enslaved by man; but, so long as he obeys a human lord, he is an auxiliary in the warfare his master is ever waging against all existences except those which he can tame to a willing servitude.
Number of Quadrupeds in the United States.
Civilization is so intimately associated with, if not dependent upon, certain inferior forms of animal life, that cultivated man has never failed to accompany himself, in all his migrations, with some of these humble attendants. The ox, the horse, the sheep, and even the comparatively useless dog and cat, as well as several species of poultry, are voluntarily transported by every emigrant colony, and they soon multiply to numbers very far exceeding those of the wild genera most nearly corresponding to them.[62] According to the census of the United States for 1860,[63] the total number of horses