forms a substance whose atoms severally contain at least four ultimate atoms of three different kinds. Yet more heterogeneous and less stable still are the salts; which present us with molecules each made up of five, six, seven, eight, ten, twelve, or more atoms, of three, if not more, kinds. Then there are the hydrated salts, of a yet greater heterogeneity, which undergo partial decomposition at much lower temperatures. After them come the further complicated supersalts and double salts, having a stability again decreased; and so throughout. Without entering into qualifications for which space fails, we believe no chemist will deny it to be a general law of these inorganic combinations that, other things equal, the stability decreases as the complexity increases. When we pass to the compounds of organic chemistry, we find this general law still further exemplified: we find much greater complexity and much less stability. A molecule of albumen, for instance, consists of 482 ultimate atoms of five different kinds. Fibrine, still more intricate in constitution, contains in each molecule, 298 atoms of carbon, 49 of nitrogen, 2 of sulphur, 228 of hydrogen, and 92 of oxygen—in all, 669 atoms; or, more strictly speaking, equivalents. And these two substances are so unstable as to decompose at quite ordinary temperatures; as that to which the outside of a joint of roast meat is exposed. Thus it is manifest that the present chemical heterogeneity of the Earth's surface has arisen by degrees, as the decrease of heat has permitted; and that it has shown itself in three forms—first, in the multiplication of chemical compounds; second, in the greater number of different elements contained in the more modern of these compounds; and third, in the higher and more varied multiples in which these more numerous elements combine.
To say that this advance in chemical heterogeneity is due to the one cause, diminution of the Earth's temperature, would be to say too much; for it is clear that aqueous and atmospheric agencies have been concerned; and further, that the affinities of the elements themselves are implied. The cause has all along been a composite one: the cooling of the Earth having been simply the most general of the concurrent causes, or assemblage of conditions. And here, indeed, it may be remarked that in the several classes of facts already dealt with (excepting, perhaps, the first), and still more in those with which we shall presently deal, the causes are more or less compound; as indeed are nearly all causes with which we are acquainted. Scarcely any change can rightly be ascribed to one agency alone, to the neglect of the permanent or temporary conditions under which only this agency produces the change. But as it does not materially affect our argument, we prefer, for simplicity's sake, to use throughout the popular mode of expression. Perhaps it will be further objected, that to assign loss of heat as the cause of any changes, is to attribute these changes not to a force, but to the absence of a force. And this is true. Strictly speaking, the changes should be attributed to those forces which come into action when the antagonist force is withdrawn. But though there is inaccuracy in saying that the freezing of water is due to the loss of its heat, no practical error arises from it; nor will a parallel laxity of expression vitiate our statements respecting the multiplication of effects. Indeed, the objection serves but to draw attention to the fact, that not only does the exertion of a force produce more than one change, but the withdrawal of a force produces more than one change.
Returning to the thread of our exposition, we have next to trace, throughout organic progress, this same all-pervading principle. And here, where the evolution of the homogeneous into the heterogeneous was first observed, the production of many effects by one cause is least easy to demonstrate. The development of a seed into a plant, or an ovum into an animal, is so gradual, while the forces which determine it are so involved, and at the same time so unobtrusive, that it is difficult to detect the multiplication of effects which is elsewhere so obvious. But, guided by indirect evidence, we may safely conclude that here too the law holds. Note, first, how numerous are the changes which any marked action works upon an adult organism—a human being, for instance. An alarming sound or sight, besides the impressions on the organs of sense and the nerves, may produce a start, a scream, a distortion of the face, a trembling consequent on general muscular relaxation, a burst of perspiration, a rush of blood to the brain, followed possibly by arrest of the heart's action and by syncope; and if the subject be feeble, an indisposition with its long train of complicated symptoms may set in. Similarly in cases of disease. A minute portion of the small-pox virus introduced into the system, will, in a severe case, cause, during the first stage, rigors, heat of skin, accelerated pulse, furred tongue, loss of appetite, thirst, epigastric uneasiness, vomiting, headache, pains in the back and limbs, muscular weakness, convulsions, delirium, &c.; in the second stage, cutaneous eruption, itching, tingling, sore throat, swelled fauces, salivation, cough, hoarseness, dyspnœa, &c.; and in the third stage, œdematous inflammations, pneumonia, pleurisy, diarrhœa, inflammation of the brain, ophthalmia, erysipelas, &c.: each of which enumerated symptoms is itself more or less complex. Medicines, special foods, better air, might in like manner be instanced as producing multiplied results. Now it needs only to consider that the many changes thus wrought by one force upon an adult organism, will be in part paralleled in an embryo organism, to understand how here also, the evolution of the homogeneous into the heterogeneous may be due to the production of many effects by one cause. The external heat, which, falling on a matter having special proclivities, determines the first complications of the germ, may, by acting on these, superinduce further complications; upon these still higher and more numerous ones; and so on continually: each organ as it is developed serving, by its actions and reactions on the rest, to initiate new complexities. The first pulsations of the fœtal heart must simultaneously aid the unfolding of every part. The growth of each tissue, by taking from the blood special proportions of elements, must modify the constitution of the blood; and so must modify the nutrition of all the other tissues. The heart's action, implying as it does a certain waste, necessitates an addition to the blood of effete matters, which must influence the rest of the system, and perhaps, as some think, cause the formation of excretory organs. The nervous connexions established among the viscera must further multiply their mutual influences; and so continually. Still stronger becomes the probability of this view when we call to mind the fact, that the same germ may be evolved into different forms according to circumstances. Thus, during its earlier stages, every embryo is sexless—becomes either male or female as the balance of forces acting on it determines. Again, it is a well-established fact that the larva of a working-bee will develop into a queen-bee, if before it is too late, its food be changed to that on which the larvæ of queen-bees are fed. All which instances suggest that the proximate cause of each advance in embryonic complication is the action of incident forces upon the complication previously existing. Indeed, we may find a priori reason to think that the evolution proceeds after this manner. For since no germ, animal or vegetal, contains the slightest rudiment or indication of the future organism—since the microscope has shown us that the first process set up in every fertilized germ, is a process of repeated spontaneous fissions ending in the production of a mass of cells, not one of which exhibits any special character; there seems no alternative but to suppose that the partial organization at any moment existing in a growing embryo, is transformed by the agencies acting upon it into the succeeding phase of organization, and this into the next, until, through ever-increasing complexities, the ultimate form is reached. Not indeed that we can thus really explain the production of any plant or animal. We are still in the dark respecting those mysterious properties in virtue of which the germ, when subject to fit influences, undergoes the special changes that begin the series of transformations. All we aim to show, is, that given a germ possessing those particular proclivities distinguishing the species to which it belongs, and the evolution of an organism from it, probably depends on that multiplication of effects which we have seen to be the cause of progress in general, so far as we have yet traced it.
When, leaving the development of single plants and animals, we pass to that of the Earth's flora and fauna, the course of our argument again becomes clear and simple. Though, as was admitted in the first part of this article, the fragmentary facts Paleontology has accumulated, do not clearly warrant us in saying that, in the lapse of geologic time, there have been evolved more heterogeneous organisms, and more heterogeneous assemblages of organisms, yet we shall now see that there must ever have been a tendency towards these results. We shall find that the production of many effects by one cause, which as already shown, has been all along increasing the physical heterogeneity of the Earth, has further involved an increasing heterogeneity in its flora and fauna, individually and collectively. An illustration will make this clear. Suppose that by a