Again, in the unmodified ectoderm, as we see it in the Hydra, the units are all endowed both with impressibility and contractility; but as we ascend to higher types of organization, the ectoderm differentiates into classes of units which divide those two functions between them: some, becoming exclusively impressible, cease to be contractile; while some, becoming exclusively contractile, cease to be impressible. Similarly with societies. In an aboriginal tribe, the directive and executive functions are diffused in a mingled form throughout the whole governing class. Each minor chief commands those under him, and, if need be, himself coerces them into obedience. The council of chiefs itself carries out on the battle-field its own decisions. The head chief not only makes laws, but administers justice with his own hands. In larger and more settled communities, however, the directive and executive agencies begin to grow distinct from each other. As fast as his duties accumulate, the head chief or king confines himself more and more to directing public affairs, and leaves the execution of his will to others: he deputes others to enforce submission, to inflict punishments, or to carry out minor acts of offence and defence; and only on occasions when, perhaps, the safety of the society and his own supremacy are at stake, does he begin to act as well as direct. As this differentiation establishes itself, the characteristics of the ruler begin to change. No longer, as in an aboriginal tribe, the strongest and most daring man, the tendency is for him to become the man of greatest cunning, foresight, and skill in the management of others; for in societies that have advanced beyond the first stage, it is chiefly such qualities that insure success in gaining supreme power, and holding it against internal and external enemies. Thus that member of the governing class who comes to be the chief directing agent, and so plays the same part that a rudimentary nervous centre does in an unfolding organism, is usually one endowed with some superiorities of nervous organization.
In those larger and more complex communities possessing, perhaps, a separate military class, a priesthood, and dispersed masses of population requiring local control, there grow up subordinate governing agents; who, as their duties accumulate, severally become more directive and less executive in their characters. And when, as commonly happens, the king begins to collect round himself advisers who aid him by communicating information, preparing subjects for his judgment, and issuing his orders; we may say that the form of organization is comparable to one very general among inferior types of animals, in which there exists a chief ganglion with a few dispersed minor ganglia under its control.
The analogies between the evolution of governmental structures in societies, and the evolution of governmental structures in living bodies, are, however, more strikingly displayed during the formation of nations by coalescence of tribes—a process already shown to be, in several respects, parallel to the development of creatures that primarily consist of many like segments. Among other points of community between the successive rings which make up the body in the lower Annulosa, is the possession of similar pairs of ganglia. These pairs of ganglia, though connected by nerves, are very incompletely dependent on any general controlling power. Hence it results that when the body is cut in two, the hinder part continues to move forward under the propulsion of its numerous legs; and that when the chain of ganglia has been divided without severing the body, the hind limbs may be seen trying to propel the body in one direction while the fore limbs are trying to propel it in another. But in the higher Annulosa, called Articulata, sundry of the anterior pairs of ganglia, besides growing larger, unite in one mass; and this great cephalic ganglion having become the co-ordinator of all the creature's movements, there no longer exists much local independence. Now may we not in the growth of a consolidated kingdom out of petty sovereignties or baronies, observe analogous changes? Like the chiefs and primitive rulers above described, feudal lords, exercising supreme power over their respective groups of retainers, discharge functions analogous to those of rudimentary nervous centres. Among these local governing centres there is, in early feudal times, very little subordination. They are in frequent antagonism; they are individually restrained chiefly by the influence of parties in their own class; and they are but irregularly subject to that most powerful member of their order who has gained the position of head-suzerain or king. As the growth and organization of the society progresses, these local directive centres fall more and more under the control of a chief directive centre. Closer commercial union between the several segments is accompanied by closer governmental union; and these minor rulers end in being little more than agents who administer, in their several localities, the laws made by the supreme ruler: just as the local ganglia above described, eventually become agents which enforce, in their respective segments, the orders of the cephalic ganglion. The parallelism holds still further. We remarked above, when speaking of the rise of aboriginal kings, that in proportion as their territories increase, they are obliged not only to perform their executive functions by deputy, but also to gather round themselves advisers to aid in their directive functions; and that thus, in place of a solitary governing unit, there grows up a group of governing units, comparable to a ganglion consisting of many cells. Let us here add that the advisers and chief officers who thus form the rudiment of a ministry, tend from the beginning to exercise some control over the ruler. By the information they give and the opinions they express, they sway his judgment and affect his commands. To this extent he is made a channel through which are communicated the directions originating with them; and in course of time, when the advice of ministers becomes the acknowledged source of his actions, the king assumes the character of an automatic centre, reflecting the impressions made on him from without.
Beyond this complication of governmental structure many societies do not progress; but in some, a further development takes place. Our own case best illustrates this further development and its further analogies. To kings and their ministries have been added, in England, other great directive centres, exercising a control which, at first small, has been gradually becoming predominant: as with the great governing ganglia which especially distinguish the highest classes of living beings. Strange as the assertion will be thought, our Houses of Parliament discharge, in the social economy, functions which are in sundry respects comparable to those discharged by the cerebral masses in a vertebrate animal. As it is in the nature of a single ganglion to be affected only by special stimuli from particular parts of the body; so it is in the nature of a single ruler to be swayed in his acts by exclusive personal or class interests. As it is in the nature of a cluster of ganglia, connected with the primary one, to convey to it a greater variety of influences from more numerous organs, and thus to make its acts conform to more numerous requirements; so it is in the nature of the subsidiary controlling powers surrounding a king to adapt his rule to a greater number of public exigencies. And as it is in the nature of those great and latest-developed ganglia which distinguish the higher animals, to interpret and combine the multiplied and varied impressions conveyed to them from all parts of the system, and to regulate the actions in such way as duly to regard them all; so it is in the nature of those great and latest-developed legislative bodies which distinguish the most advanced societies, to interpret and combine the wishes of all classes and localities, and to make laws in harmony with the general wants. We may describe the office of the brain as that of averaging the interests of life, physical, intellectual, moral; and a good brain is one in which the desires answering to these respective interests are so balanced, that the conduct they jointly dictate, sacrifices none of them. Similarly, we may describe the office of a Parliament as that of averaging the interests of the various classes in a community; and a good Parliament is one in which the parties answering to these respective interests are so balanced, that their united legislation allows to each class as much as consists with the claims of the rest. Besides being comparable in their duties, these great directive centres, social and individual, are comparable in the processes by which their duties are discharged. The cerebrum is not occupied with direct impressions from without but with the ideas of such impressions. Instead of the actual sensations produced in the body, and directly appreciated by the sensory ganglia, or primitive nervous centres, the cerebrum receives only the representations of these sensations; and its consciousness is called representative consciousness, to distinguish it from the original or presentative consciousness. Is it not significant that we have hit on the same word to distinguish the function of our House of Commons? We call it a representative body, because the interests with which it deals are not directly presented to it, but represented to it by its various members; and a debate is a conflict of representations of the results likely to follow from a proposed