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Biogeography in the Sub-Arctic


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Section I Remote Origins

       Brian G. J. Upton

       School of GeoSciences, University of Edinburgh, UK

      The northern landmasses, namely North America, Greenland and Europe/Asia were part of one global super‐continent in the lower Palaeozoic, approximately 420–430 million years (Ma) ago. This super‐continent (Pangaea) resulted from continental collisions. Driven by convective flow deep in the interior of the Earth it is the nature of continents to break apart, re‐join and to come apart again. Such a separation and amalgamation constitutes ‘the Wilson Cycle’, which takes several hundred million years to run its full course. No sooner had Pangaea come into existence than it became subject to tectonic stresses that tended to disrupt it. The North Atlantic Ocean is a late product of the disintegration of Laurasia, a part of Pangaea, which split to form North America, Greenland, Europe and Asia. Continental separation had begun in the south Atlantic region at ca 130 Ma and spread north by 60–50 Ma.

      Before considering the birth and growth of the North Atlantic, a brief résumé concerning plate tectonics is in order. Volumetrically the greater bulk of the planet is composed of the mantle. The latter, itself covered by thin veneers of crust, hydrosphere and atmosphere, extends down to a depth of 2885 km, i.e. to the outer boundary of the core from which it receives heat. The mantle is composed of various magnesium‐rich silicates and oxides and is deduced to behave as a ductile material that is in constant slow convective motion, flowing whilst remaining (almost entirely) in the solid state. The flowage is due to variations in its composition and temperature (principally the latter) that confer different densities to some parts. Consequently, the relative buoyancy of those parts with lower density causes them to rise while, simultaneously, other denser parts sink to take their place.

      Continental lithosphere differs from its oceanic counterpart not only in being older and thicker but in composition, complexity and overall lower density. The last of these causes it to ‘float’ higher so that most rises above sea‐level and the remainder is covered only by shallow seas and constitutes the continental shelves. By contrast, because of its higher density, the oceanic lithosphere lies at lower levels than its continental counterpart and its surface is almost invariably submarine, with the ocean floors generally lying at depths of ~4 km. The oceanic lithosphere, with its constructive plate boundaries (mid‐ocean ridges) and corresponding destructive plate boundaries (along subduction zones, generally demarcated by deep ocean trenches), covers some 5/7th of the Earth's surface. The continental shelves slope steeply down from depths of ~2 km to the deep ocean floors. Consequently, the submarine 2 km contour approximates the change‐over from one type of lithosphere to the other. On the western side of the North Atlantic the continental shelf is very narrow, contrasting with the European side where it is much broader.

      The ocean floor moves away symmetrically on either side of the mid‐ocean ridges as new lithosphere is generated along them. The process of ocean‐floor spreading causes a reduction of pressure along the axes which in turn promotes partial melting, to the extent of some 10–15%, of the asthenospheric mantle that underlies the lithosphere. The melt product is basaltic magma that, being less dense than the residual solid mantle, ascends towards the surface. Losing heat as it does so, most crystallizes to form intrusive rocks within the oceanic crust and the remainder erupts as lava on the ocean floor. Although the volume of lava erupted from the mid‐ocean ridges is estimated to be between 2 and 3 km3/year, a much greater volume is estimated to crystallize to form the underlying intrusions.

      The spreading process confers bilateral symmetry to the mid‐ocean ridges as juvenile material is welded on either side to the pre‐existing lithosphere. The mantle rocks that are not consumed in the melting remain behind, contributing to the lower part of the oceanic lithosphere. Sea‐floor spreading occurs at rates that can exceed 100 mm/year, whilst that in the North Atlantic is roughly 25 mm/year (Bown and White 1994).

An illustration of a map depicting the bathymetry of the North Atlantic, based on satellite sea-surface altimetry, model DNSC08. The symmetrical disposition of the mid-Atlantic ridge relative to the bounding continents is well exhibited.

      Source: Based on Anderson, O.B. & Knudsen, P. 2009.

      Because of this mechanism the spreading (and thus growth of the oceanic plate) does not take place continuously but in a jerky, spasmodic manner. For example, generation of a new fissure 1 m wide every 50 years would correspond to an averaged spreading rate of 20 mm/year. Hence,